Integrative properties associated with each behavioral state
| Behavioral state | Theta | Sharp-wave |
|---|---|---|
| Integration mode | Linear | Nonlinear |
| Integrating locus | Soma | Dendrite |
| Input arrival timing | Asynchronous (100–200 ms) | Synchronous (<20 ms) |
| Spatial distribution of inputs | Distributed | Clustered |
| Action potentials generated | Multiple | Single |
| Dendritic function | Antenna | Pattern detector |