Cover legend: Immunofluorescent labeling of the planar cell polarity molecule Prickle-like 2 (PK2; green) in pairs of vestibular hair cells with opposite stereocilia bundle polarities from saccule (top row) and utricle (bottom row). Polarity is determined by the position of a void in α-spectrin labeling (red). Note that the subcellular distribution of Pk2 protein remains constant despite the opposite polarity of cells in either pair. For more information, see the article by Deans et al. in this issue (pages 3139–3147).