Cover legend: The axons of an ON cone bipolar cell type (red with long, descending axons) contain synaptic ribbons (green) high in their axons, which make contact with dopaminergic amacrine cells (cyan) of the rabbit retina. These axonal synapses explain the ON light responses of three retinal cell types, which ramify in the OFF layer and challenge the traditional ON/OFF segregation of bipolar cell outputs. For more information, see the article by Hoshi et al. in this issue (pages 8875–8883).