Cover legend: These vasoactive intestinal peptide (VIP)-expressing interneurons in mouse somatosensory cortex were targeted with a modified rabies virus system to label their brain-wide monosynaptic inputs. Many VIP-expressing interneurons exhibit a striking bipolar morphology, with primary neurites that run perpendicular to the cortical surface. For more information, see the article by Wall et al. (pages 4000–4009).