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This composite image shows channelrhodopsin-expressing axons of the lateral perforant path (green) targeting dendritic segments of granule cells (red) in the outer molecular layer of the mouse hippocampus. Laminar-specific activation showed that newborn cells receive strong preferential input from the lateral perforant path, despite showing similar spine density and dendritic length in the middle (light blue) and outer (light purple) molecular layer, a feature that may support their unique role in pattern separation. (pages 5843–5853).